To request a reprint or PDF, please contact Dawn Potter (

* Any complete list of publications resulting from work done under my supervision should also include "publications of graduate students". This is because it has always been my philosophy not to add my name to papers resulting from research performed by my graduate students. Those publications are listed elsewhere on this web site.

118. Emlen, S. T. 2010. Taking my cues from nature's clues. Pp. 241-278, in: Leaders in Animal Behavior: The Second Generation, editors. L.C. Drickamer & D.A. Dewsbury, Cambridge University Press, Cambridge, U.K. (This book is a collection of invited autobiographies from 21 individuals who have been peer selected and have provided unique and important contributions to the field in the past 25 years.)
117. Buston, P.M., Reeve, H.K., Cant, M.A., Vehrencamp, S.L., & Emlen, Stephen T. 2007. Reproductive skew and the evolution of group dissolution tactics: a synthesis of concession and restraint models. Animal Behaviour, 74:1643-1654.  
116. Wrege, P.H. and Emlen, S.T. 2005. Sexing criteria, accuracy, and statistical inference-a reply. Auk 122:348-349.  
115. McRae, S.B., Emlen, S.T., Rubenstein, D.R., and Bogdanowicz, S.M. 2005. Polymorphic microsatellite loci in a plural breeder, the grey-capped social weaver (Pseudonigrita arnaudi), isolated with an improved enrichment protocol using fragment size-selection. Molecular Ecology Notes 5(1):16-20.  
114. Emlen, S.T. and Wrege, P.H. 2004. Size dimorphism, intra-sexual competition, and sexual selection in a sex-role-reversed shorebird, the Wattled Jacana (Janaca jacana), The Auk 121:391-403.  
113. Emlen, S.T. and Wrege, P.H. 2004. Division of labour in parental care behaviour of a sex-role-reversed shorebird, the wattled jacana. Animal Behaviour 68:847-855.  
112. Buston, P.M. and Emlen, S.T. 2003. Cognitive processes underlying human mate choice: The relationship between self-perception and mate preference in Western society. Proceedings of the National Academy of Science, 100(15):8805-8810.  
____ Subject of an article in the New York Times (Science Times) by Natalie Angier ("Opposites attract? Not in real life"), July 8, 2003.  
____ Subject of website articles on New ("Opposites do not attract in mating game" by Catherine Zandonella, June 30, 2003), ("Opposites `do not attract'", July 2, 2003); (July 2, 2003); ("Fidelity is what most people want", July 8, 2003); ABC Online Australia ("Mate choice: opposites don't attract", July 4, 2003).  
____ Subject of newspaper articles in: The Globe and Mail, Canada ("Seeking the perfect mate? Try someone just like you", by Anne McIlroy, July 1, 2003), The Boston Globe ("Like likes like" by Agnieszka Bishop, July 8, 2003), Washington Timnes ("Fidelity is what most people want", July 8, 2003); The Daily Times of Pakistan ("Opposites do not attract", July 1, 2003), The Times of India ("Do opposites attract?", July 9, 2003 and "Not everyone wants to marry a millionaire" by Vithal Nadkarni, July 5, 2003), The Western Mail, Wales ("Opposites don't always attract" by Duncan Higgitt, July 1, 2003), International Herald Tribune ("Seeking a spouse? Take a look in the mirror, new study says", July 7, 2003); Daily Telegraph, Australia ("Like stays with like", July 1, 2003), UPI International; Reuters and many others...  
____ Subject of interviews on National Public Radio, USA (the Todd Mundt Show); BBC World Service; BBC Late Show; BBC Live at Five; News Center 10; Bloomberg News; News Talk 106 (Ireland); Bruce Kenyan Show (Canada), and others...  
111. Emlen, S.T. 2001. Introduction to William D. Hamilton Symposium. Behavioral Ecology 12:261.  
110. Emlen, S.T. and Demong, N.J. 2001. Bird families as models for understanding ourselves. Cornell Laboratory of Ornithology, Homestudy Course in Bird Biology, Chapter 6, Understanding Bird Behavior, pp. 91-94.  
109. Reeve, H.K. and Emlen, S.T. 2000. Reproductive skew and group size: an N-person staying incentive model. Behavioral Ecology 11: 640-647.
108. Whittingham, L.A., Sheldon, F.H., and Emlen, S.T. 2000. Molecular phylogeny of jacanas and its implications for morphologic and biogeographic evolution.. Auk, 117(1):22-32.
107. Lanyon, W.E., Emlen, S.T., and Orians, G.H. 2000. In Memoriam: John Thompson Emlen, Jr., 1908-1997. Auk, 117(1):222-227.
106. Emlen, S.T. 1999. Reproductive skew in cooperatively breeding birds: an overview of the issues, pp. 2922-2931, In: Adams, N.J. & Slotow, R.H. (eds) Proc. 22nd International Ornithological Congress. Johannesburg: BirdLife South Africa.
105. du Plessis, M. and Emlen, S.T. (eds.) 1999. The sociality continuum: monopolized and shared breeding systems among cooperatively breeding birds, pp. 2857-2931 (Symposium No. 47), In: Adams, N.J. & Slotow, R.H. (eds) Proc. 22nd International Ornithological Congress. Johannesburg: BirdLife South Africa.
104. Kaplan, D. M. and Vanduser, M. L. 1999. Evolution and Stepfamilies: An Interview with Dr. Stephen T. Emlen, The Family Journal, Vol. 7:408-413. (The Family Journal is the official publication of the International Association of Marriage and Family Counselors), (October 1999).
103. Emlen, S.T. 1999. Foreword to the 1999 Edition of Helpers at Birds' Nests, by Alexander F. Skutch, pp. ix-xii, University of Iowa Press, Iowa City.  
102. Emlen, S.T. 1998. Relevance and responsibility in Behavioural Ecology. Presidential address to the International Society for Behavioural Ecology. ISBE Newsletter, 10(2):8-10.
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101. Emlen, S.T., Wrege, P.H., Webster, M.S. 1998. Cuckoldry as a cost of polyandry in the sex-role reversed wattled jacana, Jacana jacana, Proceedings of the Royal Society of London, Series B, 265:2359-2364
____ (Subject of an article in New York Times (Science Times) by Carol Kaesuk Yoon, ("In This Battle of the Sexes, the Females Win"), Tuesday, December 22, 1998).  
____ (Subject of an article in ScienceNOW by C. Ferguson, ("The feathered feminist"), December 22, 1998.)  
____ (Subject of note in The Sciences (The Magazine of the New York Academy of Sciences), Vol 39(2):8, March/April 1999.  
____ (Subject of an article in Birder's World ("In Battle of Sexes, Female's Triumph"), Vol 13(2):12, April 1999.)  
____ (Subject of an article in International Wildlife Magazine by Les Line, ("A male's work is never done"), Vol. 30(4):28-35. July/August 2000.  
100. Emlen, S.T., Reeve, H.K., & Keller, L. 1998. Reproductive skew: disentangling concessions from control. Trends in Ecology and Evolution, 13: 458-459.  
99. Reeve, H.K., Emlen, S.T., and Keller, L. 1998. Reproductive sharing in animal societies: reproductive incentives or incomplete reproductive control by dominant breeders? Behavioural Ecology, 9(3):267-276.
98. Emlen, S.T. 1997. The evolutionary study of human family systems. Social Science Information 36: 563-589.
____ (Subject of article in Conocer Magazine (Spain) by Merideth Small ("It's all in the family") (Todo queda en la familia,) pp. 63-70, September, 1997).  
____ (Subject of a "profile" article in Discover Magazine by Will Hiveley ("Family man"), pp. 80-90, October, 1997).  
____ (Subject of an article in The New York Times (Science Times) by Jane E. Brodie, ("Genetic Ties May Be Factor in Violence in Stepfamilies"), Tuesday, February 10, 1998)).  
____ (Subject of an article in Family Systems by D.V. Papero, ("Stephen Emlen: The Biological Basis of the Family"), Family Systems, Vol. 4(2): 185-192. (Family Systems is a professional journal of natural systems thinking in psychiatry and the sciences, published by the Georgetown Family Center in Washington, D.C.). (November 1998).  
____ (Subject of an article in The Family Journal by D.M. Kaplan and M.L. Vanduser (1999). ("Evolution and Stepfamilies: An interivew with Dr. Stephen T. Emlen"), The Family Journal, Vol. 7(4):408-413, (October 1999). (The Family Journal is the official publication of the International Association of Marriage and Family Counselors).  
97. Emlen, S.T. 1997. Human family dynamics: an evolutionary perspective. Kagaku 67: 279-288. (Translated into Japanese; Kaguku is a Japanese Scientific American-type journal).  
96. Emlen, S.T. 1997. Predicting family dynamics in social vertebrates. Review chapter for Behavioural Ecology: An Evolutionary Approach, 4th Edition. J. R. Krebs and N. B. Davies, eds., pp 228-253. Blackwell Scientific, Oxford.
95. Emlen, S.T. 1997. When mothers prefer daughters over sons. Trends in Ecology and Evolution, 12: 291-292.  
94. Emlen, S.T. 1996. Reproductive sharing in different types of kin associations. American Naturalist, 148: 756-763.
93. Emlen, S.T. I996. Living with relatives: lessons from avian family systems. Acta XXI Congressus Internationalis Ornithologici: 87-100.
____ (also published in: Ibis 138: 87-100 (1996)).
92. Demong, N.J. and Emlen, S.T. 1996. All in the family. Living Bird Quarterly (Cornell Laboratory of Ornithology) vol. 15 (Summer): 30-34.  
____ (Reprinted by invitation on the SIRS electronic database (a subscriber network distributing materials to schools and libraries in the United States and Canada.) March 1997)  
91. Emlen, S. T. 1996. Historical perspectives on orientation: an addendum. Condor 98:176.
____ Hrdy, S. B., Rodman, P., Charnov, E. L., Seger, J., Hawkes K., Emlen, S. T., Foster, S. A., Gowaty, P. A., Haig, D., Hauser, M. Jacobs, L.F., and Smuts, B. B. 1996. Sociobiology's successes. Science 274:162-163.
90. Emlen, S.T. 1995. An evolutionary theory of the family. Proceedings of the National Academy of Science (Review Article). PNAS 92(18):8092-8099.
____ (Subject of article in The New York Times (Science Times) by Natalie Angier ("New view of family: unstable but wealth helps"), August 29, 1995).  
____ (Reprinted by invitation on the electronic server of the Human Behavior and Evolution Society, 1996;
89. Emlen, S.T., P.H. Wrege and N.J. Demong 1995. Making decisions in the family: an evolutionary perspective. American Scientist 83:148-157.  
____ (Reprinted in Exploring Animal Behaviour: Readings from American Scientist, P. W. Sherman and J. Alcock, eds., Sinauer Associates, 2nd Edition, 1997).  
88. Emlen, S.T. 1995. Can avian biology be useful to the social sciences. (Invited "Point of View"), J. Avian Biology, 26: 273-276.  
87. Demong, N.J. and S.T. Emlen 1995. Signals of the flesh. Nature Australia , 25(2):26-28.  
86. Emlen, S.T. 1994. Benefits, constraints and the evolution of the family. Trends in Ecology and Evolution, (Cover Story) 9:282-285.
85. Emlen, S.T. and P.H. Wrege 1994. Gender, status and family fortunes in the White-fronted Bee-eater. Nature, (Progress Article) 367:129-132.
____ (Subject of invited article in La Repubblica (Rome) by Natalie J. Demong ("Kinship, gender and social structure are important determinants of avian family dynamics"), May 20, 1995.  
84. Wrege, P.H. and S.T. Emlen 1994. Family structure influences mate choice in White-fronted Bee-eaters. Behavioral Ecology and Sociobiology 35:185-191.
____ Fivizzani, A., Delehanty, D., Oring, L., Wrege, P., and Emlen, S.T. 1994. Elevated female androstenedione levels in two sex-role reversed avian species. American Zoologist, 34:22a.  
83. Emlen, S.T. 1993. Ethics and Experimentation: hard choices for the field ornithologist. The Auk, 110:406-409.  
82. Murphy, C.G., Emlen, S.T. and Sherman, P.W. 1993. Reproductive strategies of the treefrog, Hyla gratiosa: Implications for management. Florida Game and Fresh Water Fish Commission, final report project NG88-020.
81. Emlen, S.T. and P.H. Wrege 1992. Parent-offspring conflict and the recruitment of helpers among bee-eaters. Nature , (Cover story) 356:331-333.
____ (Subject of article in The New York Times (Science Times) by Natalie Angier ("Biologists tell a tale of interfering in-laws"), April 7, 1992).  
____ (Subject of article in National Wildlife Magazine by Bruce Fellman ("Looking out for number one"), pp. 46-49, Dec. 1992).  
____ (Subject of note in BBC Wildlife Magazine by Stephen Young ("The generation gamble"), Vol. 10, pg 13; May 1992).  
____ Subject of article in Down to Earth Magazine, (India) by R. Gadagkar, ("When Fathers Harrass Their Sons") Vol. 1 (8):47-48, September 1992.)  
____ (Subject of note in Geographica, National Geographic Magazine, Feb. 1993.)
80. Emlen, S.T. and P.H. Wrege 1992. Parent-offspring conflict. Nature , 359:24.
79. Emlen, S.T. 1992. Family dynamics in an evolutionary perspective. Video tape of Distinguished Lecture, available from Georgetown Family Center, Georgetown University, Washington, DC.  
78. Emlen, S.T. 1992. The evolution of the family. Video tape of Keynote lecture, available from Georgetown Family Center, Georgetown University, Washington, DC.  
77. Demong, N.J. and S.T. Emlen 1992. Family ties: helping in African bee-eaters. Living Bird Quarterly, (Cornell Laboratory of Ornithology) 2:26-31.  
76. Emlen, S.T. 1991. The evolution of cooperative breeding in birds and mammals. In: Behavioural Ecology: An Evolutionary Approach, 3rd Edition. J. Krebs and N. B. Davies, eds. Blackwell Scientific, Oxford, pp. 301-337.
75. Emlen, S.T., F.L.W. Ratnieks, H.K. Reeve, J. Shellman-Reeve, P.W. Sherman and P.H. Wrege 1991. Adaptive versus nonadaptive explanations of behavior: the case of alloparental helping. American Naturalist, 138:259-270.  
74. Emlen, S.T. and P.H. Wrege 1991. Breeding biology of White-fronted Bee-eaters at Nakuru: the influence of helpers on breeder fitness. Journal of Animal Ecology, 60:309-326.
73. Emlen, S.T., P.H. Wrege, N.J. Demong and R.E. Hegner 1991. Flexible growth rates in nestling White-fronted Bee-eaters: a possible adaptation to short-term food shortage. Condor, 93:591-597.
72. Wrege, P.H. and S.T. Emlen 1991. Breeding seasonality and reproductive success of White-fronted Bee-eaters in Kenya. The Auk, 108:673-687.
71. Emlen, S.T. 1990. The White-fronted Bee-eater: helping in a colonially nesting species. In: Cooperative Breeding in Birds: Long-Term Studies of Ecology and Behavior. P. Stacey and W. Koenig, eds. Cambridge University Press, Cambridge, pp. 487-526.  
____ (subject of article in International Wildlife Magazine by Les Line. ("A bug's worst nightmare,"), July-August, 1993. pp.30-37.)  
70. Emlen, S.T. 1990. Observations on a captive colony of Quaker parakeets. AFA Watchbird, 27:26-29 (Magazine of the American Federation of Aviculture).  
69. Emlen, S.T., N.J. Demong and D. Emlen 1989. Experimental induction of infanticide in female wattled jacanas. The Auk, 105:1-7.
____ (Featured in article in Discover Magazine by Jared Diamond ("Reversal of Fortune"), April 1992, pp. 70-75).  
____ (Research featured in: "The Trials of Life" ("Continuing the Line" segment), a 13 part television documentary on animal behavior produced by the BBC with Sir David Attenborough, 1993).  
____ (Featured in article in BioScience by Kathryn Brown ("Infanticide as a way to get ahead"), BioScience , 46 (3):174-177 plus inside cover, 1996).  
____ (Featured in article in Discover Magazine by Carl Zimmer ("First, kill the babies"), September 1996, pp. 73-78.)  
68. Emlen, S.T. and P.H. Wrege 1989. A test of alternate hypotheses for helping behavior in White-fronted Bee-eaters of Kenya. Behavioral Ecology and Sociobiology, 25:303-320.  
____ (Subject of article in Current Science , ("India"), by R. Gadagkar and A.B. Venkataraman ("Nepotistic Bee-eaters"), Volume 59:445-446, 1990).  
67. Konishi, M., S.T. Emlen, R.E. Ricklefs and J.C. Wingfield 1989. Contributions of bird studies to biology. Science, 246:465-472.
____ (Reprinted in: Biological Systems, B.R. Jasnt and D.E. Koshland, Jr., eds., American Association Advancement of Science, Wash. D.C., 1990).  
66. Emlen, S.T. and P.H. Wrege 1988. The role of kinship in helping decisions among White-fronted Bee-eaters. Behavioral Ecology and Sociobiology, 23:305-315.
____ (Subject of article in BBC Wildlife Magazine [Cover story] by A. Fothergill ("Neighbors"), Volume 6:142-148, 1988.)
____ (Subject of article in Science News [Cover story] by Kathy A. Fackelmann ("Avian Altruism"), Volume 135:364-365, 1989.)  
____ (Subject of note in The Chronical of Higher Education, Jan 11, 1989, p. A5).
____ (Featured in article in Mosiac Magazine [publication of the National Science Foundation] by A. Fisher ("A new synthesis comes of age: How different are humans?"), Volume 22(1):10-17, 1991).  
65. Emlen, S.T. and P.H. Wrege 1988. The Bee Team. BBC Documentary (Produced by Alistair Fothergill; filmed by Simon King). Shown in the Europe in the "WILDLIFE ON ONE" program and in the USA in the PBS "NATURE" Series.
____ (Winner: 7th Annual Outstanding Film Award, Animal Behavior Society, 1990).  
64. Emlen, S.T. and L.W. Oring 1988. Ecology, sexual selection and the evolution of mating systems. Citation Classic, Current Contents 18:12.  
63. Hegner, R.E. and S.T. Emlen 1987. Territorial organization of the White-fronted Bee-eater in Kenya. Ethology, 76:189-222.  
62. Wrege, P.H. and S.T. Emlen 1987. Biochemical determination of parental uncertainty in White-fronted Bee-eaters. Behavioral Ecology and Sociobiology, 20:153-160.
61. Emlen, S.T., J.M. Emlen and S.A. Levin 1986. Sex-ratio selection in species with helpers-at-the-nest. American Naturalist, 127:1-8.
60. Emlen, S.T. and P.H. Wrege 1986. Forced copulations and intra-specific parasitism: two costs of social living in the White-fronted Bee-eater. Ethology, 71:2-29.
____ (Subject of earlier article in Science Digest by Rachel Wilder ("Avian Peyton Place"), pgs. 105, 112, Oct. 1983).  
59. Emlen, S.T. and S.L. Vehrencamp 1985. Cooperative breeding strategies in birds. In: Experimental Behavioural Ecology and Sociobiology. M. Lindauer and B. Holldobler, eds. Fortschritte der Zoologie, pp. 359-374.
58. Emlen, S.T. and N.J. Demong 1985. Cé anche il parassita altrista. Atlante Magazine (Rome) 26:102-109.
57. Emlen, S.T. 1984. Cooperative breeding in birds and mammals. In: Behavioural Ecology: an evolutionary approach, 2nd edition. J. Krebs and N. Davies, eds. Blackwell Scientific Publishers, Oxford, pp. 305-335.
56. Emlen, S.T. and N.J. Demong 1984. Bee-eaters of Baharini. Natural History Magazine, (Cover story) 10:50-59.  
55. Brown, I., T. Larkin and S.T. Emlen 1984. Fourteen years of pigeon homing data. The Auk, 101:622-623.  
54. Emlen, S.T. and S.L. Vehrencamp 1983. Cooperative breeding strategies among birds. In: Perspectives in Ornithology. A. H. Brush and J. G. A. Clark, eds. Cambridge University Press, Cambridge, pp. 93-133.  
53. Emlen, S.T. 1982. The evolution of helping. I. An ecological constraints model. American Naturalist , 119:29-39.
52. Emlen, S.T. 1982. The evolution of helping. II. The role of behavioral conflict. American Naturalist, 119:40-53.
____ (Both articles subject of article in Nature [News and Views] by N.B. Davies (Cooperation and conflict in breeding groups) Volume 296:702-703, 1982).  
51. Hegner, R.E., S.T. Emlen and N.J. Demong 1982. Spatial organization of the White-fronted Bee-eater. Nature (Cover story) 296:264-266.
____ (Subject of article in The London Times ("Clan life remains strictly for the birds"), July 21, 1982).  
50. Emlen, S.T. 1982. The cooperative breeding system of African bee-eaters. National Geographic Research Reports, 14:167-170.  
49. Emlen, S.T. 1981. The ornithological roots of sociobiology. The Auk, 98:400-403.  
48. Emlen, S.T. 1981. Altruism, kinship, and reciprocity in the White-fronted Bee-eater. In: Natural Selection and Social Behavior. R. D. Alexander and D. Tinkle, eds. Chiron Press, pp. 217-230.  
____ (Featured in earlier article in Mosiac Magazine [publication of the National Science Foundation] by J. Pfeifer ("Seeking Human Origins: Cultural evolution"), Volume10(2):30-38, 1979).  
____ (Subject of article in Naturwisenschaftliche Rundschau (Germany) by G. Breuer ("Baby-sitting" auf Gegenseitigkeit beim Bienenfresser), Vol 35, p. 339, 1982).  
47. Emlen, S.T. 1981. In Memoriam: William T. Keeton. The Auk, 98:167-172.  
46. Emlen, S.T. 1980. Ecological determinism and sociobiology. In: Sociobiology: beyond Nature/Nurture. G. W. Barlow and J. Silverberg, eds. American Association for the Advancement of Science, selected symposium 35:125-150, Westview Press, Boulder, CO.
____ Emlen, S.T. and W. Wiltschko. 1980. (Symposium convenors and editors), Orientation in migratory birds. Proceedings XVII International Ornithological Congress. Berlin, pp. 528-565.  
45. Emlen, S.T. and N.J. Demong 1980. Decision making by nocturnal bird migrants: the integration of multiple cues. In: Proceedings XVII International Ornithological Congress. Berlin, pp. 553-560.  
44. Emlen, S.T., N.J. Demong and R.E. Hegner 1980. Bee-eaters: an alternative route to cooperative breeding? In: Proceedings XVII International Ornithological Congress. Berlin, pp. 895-901.  
43. Harvey, P.H., Emlen, S.T., Linsenmair, K.E., Markl, H., Marler, P., Manzel, E.W., Michener, C.D., Milinski, M., and Trillmich, F. 1980. Mechanisms of kin-correlated behavior. In: Evolution of Social Behavior: Hypotheses and Empirical Tests. H. Markl, ed. Dahlem Konferenzen, pp. 183-202.  
42. Wiltschko, W., R. Wiltschko, S.T. Emlen and N.J. Demong 1980. Nocturnal activity and orientation behavior during spring migration and early summer in indigo buntings, Passerina cyanea. In: Journal of Comparative Physiology, pp. 47-49.
41. Emlen, S.T. 1979. Predatory interactions between White-fronted Bee-eaters and fiscal shrikes. Scopus, 3:101-102. (Scopus is the ornithological journal of East Africa).  
40. Hegner, R.E., S.T. Emlen, N.J. Demong and C.E. Miller 1979. Helping at the nest in the White-fronted Bee-eater. Scopus, 3:9-13. (Scopus is the ornithological journal of East Africa).  
39. Rowley, I., S.T. Emlen, A.J. Gaston and G.E. Woolfenden 1979. A definition of "group". Ibis, 121:131.  
38. Emlen, S.T. 1978. The evolution of cooperative breeding in birds. In: Behavioural Ecology: an evolutionary approach. J.R. Krebs and N.B. Davies, eds. Blackwell Scientific Publications, Oxford, pp. 245-281.  
37. Demong, N.J. and S.T. Emlen 1978. Radar tracking of artificially released migrants: a new technique allowing orientational manipulation of free-flying birds. Bird Banding, 49:342-359.
36. Emlen, S.T. and N.J. Demong 1978. Orientation strategies used by free-flying migrants: a radar tracking study. In: Animal Migration, Navigation and Homing. K. Schmidt-Koenig and W.T. Keeton, eds. Springer-Verlag, pp. 283-294.
____ (Subject of earlier article in The New York Times, ("Bird flight cues studied by radar"), Aug. 20, 1972).  
____ (Subject of earlier article in Technology Review, ("Birds as navigators"), March/April 1972, pgs. 56-57).  
____ (Featured in Smithsonian Magazine article by David R. Zimmerman ("Probing mysteries of how birds can navigate the skies"), June 1979, pp. 52-60.)
____ (Research featured in: "On a Wing and a Prayer", a one-hour television documentary produced for PBS "New Explorers" Series and first shown in 1996.)  
35. Emlen, S.T. and L.W. Oring 1977. Ecology, sexual selection, and the evolution of mating systems. Science, 197:215-223.
____ (Featured in article in Science Magazine, [Research News] by Gina B. Kolata ("Sexual dimorphism and mating systems: How did they evolve?), Volume195:382-383, 1977).  
____ (Topic of invited "Citation Classic" in Current Contents describing the history of the 1977 paper, Current Contents 18:12, 1988).  
____ (Reprinted in: A Population Biology Reader: Critical Source Papers, G. Lovett-Doust, J.L. Lovett-Doust, and I.M. Weis, eds., William M.C. Brown, Publishers, 1992).  
34. Emlen, S.T. 1977. Double clutching and its significance in the bullfrog. Copeia, 1977:749-751.
33. Emlen, S.T. 1977. Cooperative breeding in the White-fronted Bee-eater. Kenya Wildlife Club Bulletin, (Cover story) 1977:8-11.
32. Emlen, S.T. 1976. An alternative case for sociobiology. Science, 192:736-738.
____ (Reprinted in: The Sociobiology Controversy, A.L. Caplan, ed., Harper and Row, Publishers, 1978).  
____ (Reprinted in: Selected Readings in Sociobiology, J.H. Hunt, ed., McGraw Hill Publishers, 1979).  
31. Emlen, S.T., W. Wiltschko, N.J. Demong, R. Wiltschko and S. Bergman 1976. Magnetic direction finding: evidence for its use in migratory indigo buntings. Science, 193:505-508.
30. Emlen, S.T. 1976. Lek organization and mating strategies in the bullfrog. Behavioral Ecology and Sociobiology, 1:283-313.
29. Emlen, S.T. 1976. Altruism in mountain bluebirds? Science, 191:808-809.  
28. Emlen, S.T. 1975. The stellar-orientation system of a migratory bird. Scientific American, (Cover story) 233:102-111.  
____ (Reprinted in: Readings from Animal Behaviour, T. Eisner and E.O. Wilson, eds., Freeman Press, 1979).
____ (Reprinted in: "Birds" "Readings from Scientific American", B.W. Wilson, ed., Freeman Press, 1980).  
____ (Research featured in "Bird Brain", a one-hour television documentary on bird navigation produced for the PBS "NOVA" series.)  
27. Emlen, S.T. and N.J. Demong 1975. Adaptive significance of synchronized breeding in a colonial bird: a new hypothesis. Science, 188:1029-1031.
____ (Selected for discussion in Science Magazine [Research News] by Gina B. Kolata ("Sociobiology (II): The Evolution of Social Systems"), Volume 156-157, 1975).  
26. Emlen, S.T., J.D. Rising and W.L. Thompson 1975. A behavioral and morphological study of sympatry in the indigo bunting and the lazuli buntings of the Great Plains. Wilson Bulletin, 87:145-179.
____ (Awarded the Ernest P. Edwards Prize [for best paper published in 1995] by the Wilson Ornithological Society).  
25. Emlen, S.T. 1975. Migration: orientation and navigation. In: Avian biology Vol. 5, D.S. Farner and J.R. King, eds. Academic Press, pp. 129-219.  
____ (Featured in article in National Geographic Magazine by Allan C. Fisher ("Mysteries of bird migration"), Volume 156(2):154-193, August 1978).  
____ (Featured in article in Time/Life Nature Science Annual by Peter Wood ("How Birds Find Their Way"), 1978, pg. 46-55).  
____ (Featured in article in Newsweek Magazine, by Sharon Begley ("The Migration Enigma"), October 15, 1979, pp. 111-116).  
24. Demong, N.J. and S.T. Emlen 1975. An optical scope for examining nest contents of tunnel-nesting birds. Wilson Bulletin , 87:550-551.  
23. Windsor, D. and S.T. Emlen 1975. Predatory-prey interactions of adult and prefledgling bank swallows and American kesterels. The Condor , 77:359-361.
22. Emlen, S.T. 1974. Problems identifying bird species by radar signature analyses: intra-specific variability. In: Proceedings of the Conference on Biological Aspects of the Bird/Aircraft Collision Problem, Clemson, SC. S. Gauthreaux, (ed.), pp. 509-524.
21. Emlen, S.T. 1972. An experimental analysis of the parameters of bird song eliciting species recognition. Behaviour, 41:130-171.
____ (Research featured in: "Why Birds Sing?", a one-hour television documentary produced for the PBS "NOVA" Series, 1984).  
20. Emlen, S.T. 1972. The ontogenetic development of orientation capabilities. In: Animal orientation and navigation. S.R. Galler et al., eds. NASA, pp. 191-210.
19. Emlen, S.T. 1972. Birds. Psychology Today, (an article on migratory bird navigation) (Cover story) 5:52-56.  
18. Emlen, S.T. 1972. Animal migrations. Chapter 12 in: The Marvels of Animal Behavior. National Geographic Society, Washington, D.C., pp. 270-285.  
17. Emlen, S.T. 1971. Celestial rotation and stellar orientation in migratory warblers. Science, 173:460-461.  
16. Emlen, S.T. 1971. The role of song in individual recognition in the indigo bunting. Zeitschrift fur Tierpsychologie, 28:241-246.
15. Emlen, S.T. 1971. Geographic variation in indigo bunting song. Animal Behaviour, 19:408-409.  
14. Emlen, S.T. 1970. Celestial rotation: its importance in the development of migratory orientation. Science, 170:1198-1201.
____ (Selected for discussion in in New Scientist ("Can birds learn to spot the celestial pole?"), Vol 24: 541, Dec. 24, 1970).  
____ (Selected for article in Science World 23:4-6, 1971).  
____ (Reprinted in: Biological Boundaries of Learning, M.E.P. Seligman and J.L. Hager, eds., Appleton-Century and Crofts Publishers, 1972).  
____ (Reprinted in: Readings in Animal Behaviour, T.E. McGill, ed., Holt, Rinehart and Winston Publishers, 1977).  
____ (Research featured in: "On a Wing and a Prayer", a one-hour television documentary produced for PBS "New Explorers" Series and first shown in 1996.)  
____ (Selected for article in Birder's World. ("Those amazing birds: a star guide," by Eldon Greij), p. 74, April, 1998).  
13. Emlen, S.T. 1970. The influence of magnetic information on the orientation of the indigo bunting, Passerina cyanea. Animal Behaviour 18:215-224.  
12. Emlen, S.T. and I. H. W. Ambrose 1970. Feeding interactions of snowy egrets and red-breasted mergansers. The Auk 87:164-165.  
11. Emlen, S.T. 1969. Bird migration: influence of physiological state upon celestial orientation. Science 165:716-718.
____ (Selected for discussion in New Scientist ("How buntings interpret the stars"), Vol.23: 414, Aug. 28, 1969).  
____ (Subject of article in Time Magazine ("Beacon for buntings"), Nov. 14, 1969).  
____ (Subject of article in Science Digest ("Birds migrate by the stars"), Oct. 1969, pgs. 43-4).  
____ (Featured in later article in The New York Times (Science Times), by Bayard Webster ("Mysteries of bird migration unravelled"), April 15, 1980.  
____ (Reprinted in: Foundations of Animal Behavior, L. Houck and L.C. Drickamer, eds., University of Chicago Press, 1996.)  
10. Emlen, S.T. 1969. The development of migratory orientation in young indigo bunting, Passerina cyanea. Living Bird 8:113-126.  


Emlen, S.T. 1969. The celestial guidance system of a migratory bird. (Cover story.) Sky and Telescope 38:3-6.  
8. Emlen, S.T. 1969. Homing ability and orientation in the painted turtle, Chrysemys picta marginata. Behaviour 33:58-76.
7. Emlen, S.T. 1968. Territoriality in the bullfrog, Rana catesbeiana. Copeia 1968:240-243.  
____ (Reprinted in: Readings in Animal Behavior, T.E. McGill, ed., Holt, Rinehart, and Winston Publishers, 1973; 3rd edition, 1977).
6. Emlen, S.T. 1968. A technique for marking anuran amphibians for behavioral study. Herpetologica 24:172-173.  
5. Emlen, S.T. 1967. Migratory orientation of the indigo bunting, Passerina cyanea. Part I. Evidence for use of celestial cues. The Auk 84:309-342.
4. Emlen, S.T. 1967. Migratory orientation of the indigo bunting, Passerina cyanea. Part II. Mechanism of celestial orientation. The Auk 84:463-482.
____ (Both papers subject of article in Science News by P. McBroom ("North Sky Guides Birds"), volume 90, page 152).  
____ (Subject of article in The London Times ("Bird migration may rely on constellations"), Aug. 1, 1966).  
3. Emlen, S.T. 1967. Orientation of Zugunruhe in the rose-breasted grosbeak, Pheuticus ludovicianus. Condor 69:203-205.  
2. Emlen, S.T. and J.T. Emlen, Jr. 1966. A technique for recording migratory orientation of captive birds. The Auk 83:361-367.
1. Emlen, S.T. and W. Kem 1963. Activity rhythm in Peromyscus: its influence on rates of recovery from nembutol. Science 145:1682-1683.